Born August 13, 1918, Sanger first won the prize in 1958 for his work on the structure of proteins, notably insulin, and again shared the coveted prize with two others in 1980 for pioneering developments in DNA sequencing that are still being used today, media reported.

He died on Tuesday morning at Addenbrooke's Hospital in Cambridge, eastern England, according to a spokesman for the Medical Research Council.

His colleagues described him as an inspiration: a doggedly determined but self-effacing scientist whose contribution to modern genetics and molecular biology is impossible to overstate.


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